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Silviculture8 chapters · 16 parts

Tree and forest, locality factors, succession, forest types, natural and artificial regeneration, tending.

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All chapters
Chapter 2The Tree and the ForestPart 1Part 2
Chapter 3Factors of LocalityPart 1Part 2
Chapter 4Plant SuccessionPart 1Part 2
Chapter 5Forest TypesPart 1Part 2
Chapter 6Natural RegenerationPart 1Part 2
Chapter 7Artificial RegenerationPart 1Part 2: NurseryPart 3: Planting
Chapter 8Tending OperationsPart 1Part 2
Agroforestry8 chapters · 17 parts

Concepts, classification, taungya, AF systems, tree–crop interactions, D&D, MPTs and important AF trees.

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All chapters
Chapter 1Introduction to AgroforestryPart 1Part 2
Chapter 4AF Systems and PracticesPart 1Part 2Part 3
Chapter 5Interactions, Soil and N-fixationPart 1Part 2
Chapter 6Planning, D&D and EvaluationPart 1Part 2
Chapter 7MPTs, Management and NurseryPart 1Part 2
Chapter 8AF Tree SpeciesPart 1Part 2Part 3Part 4
Forest Utilisation19 chapters

Felling, logging and transport, wood structure and properties, seasoning, preservation, MFP, pulp and paper.

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More quick revision: Forestry One Liner 123Indigenous pinesAF systems around the globe

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Forest Utilisation Chapter 8: Anatomical Structure of Wood, Sapwood, Heartwood, Vessels and Rays

Forest Utilisation Notes · Chapter 8

Chapter 8: Anatomical Structure of Wood

Cell development and pits, pith, sapwood and heartwood, bark, early and late wood, growth rings, vessels, ring and diffuse porous woods, tyloses, tracheids, fibres, parenchyma, rays, ripple marks and resin canals.

1. The cell

A group of similar cells doing the same work is a tissue. Height growth takes place at the tips and girth growth in the cambium.

Cell division
↓
Cell enlargement and wall thickening
↓
Lignification (cell hardening)
PitsDuring the course of secondary thickening certain places in the primary cell wall are left uncovered, and these thin places in the wall which serve as a means of inter-communication between two adjacent cells are known as pits.

Liquids move through the tree mainly through pits. Once lignified, a cell cannot enlarge further.

2. Gross structure (seen with naked eye or pocket lens)

Pith

Small soft mass of tissue in the centre; no special function and no diagnostic value.

Sapwood

Light coloured outer part with the youngest rings; food storage and sap conduction. Liable to fungal attack because of stored starch.

Heartwood

Deep coloured central dead tissue giving mechanical support; more durable because of gums, resins and oils that are toxic to fungi and insects.

Bark

Outer corky dead zone and inner living zone that conducts and stores food; useful in identification.

Sapwood and heartwoodThe light coloured outer portion of a tree log consisting of the youngest growth rings is called sapwood and the deep coloured central portion heartwood.
Growth ringsThese are concentric rings which usually represent the wood formed during one growing season; also called, though less aptly, annual rings.

'All sapwood' trees show no colour difference between sapwood and heartwood: fir, spruce, mango, haldu, semal, bahera, Ficus, bel and kadam. Colour can mislead: in ebony the black wood is not true heartwood, and sal sapwood can be reddish.

Early (spring) wood is formed fast with thin-walled cells and more pores; late (summer or autumn) wood is slower, thick-walled and darker. Rings are 5 to 25 mm wide in fast-grown timber and 8 or more per cm in slow-grown timber.

3. Minute structure

ElementDescription
Vessels (pores)Short wide cells forming tubes that conduct sap in broad-leaved trees. Absent in conifers except near the pith.
TracheidsIn softwoods, one cell type does both conduction and support: hollow, needle-shaped, with large bordered pits; cross-section like a honeycomb.
FibresNarrow spindle-shaped cells with pointed closed ends; give strength in broad-leaved woods.
ParenchymaShort thin-walled cells for storage and conduction of food; vertical parenchyma usually absent in conifers.
Wood (medullary) raysHorizontal cells running radially; radial conduction and food storage. Give silver grain on quarter-sawn faces.
TylosesIngrowths into vessels, foam-like, mostly in heartwood. Increase durability but make impregnation difficult.
Pith flecksPatches of parenchyma caused by insects in the cambium
Ripple marksRays in tiers (storeys) giving fine wavy lines on the tangential face; useful for identification
Resin canals and gum ductsIntercellular cavities. Pine and spruce have vertical and horizontal canals; sal and toon have gum ducts in bands; gurjan, chir and spruce have them evenly spread.
Ring-porous wood

Early-wood vessels clearly larger than late-wood vessels, forming a band. Rare in India: teak and toon.

Diffuse-porous wood

Vessels of similar size scattered through the ring. The majority of Indian hardwoods, owing to a long growing season.

Memory trick: Teak and Toon are Two ring-porous Timbers. Most Indian hardwoods are diffuse-porous.
Exam point: On the transverse section rays appear as radial lines, on the tangential section as short vertical lines, and on the radial section as horizontal ribbons.

4. Shape and size of trees

  • Species: fir and pine give tall straight stems; mango and oaks branch in the open.
  • Density of crop: dense crops give long clean boles and small crowns.
  • Age: near maturity branchwood is only 8 to 10% of the stem wood.
  • Soil and locality: better sites give more stem wood.
  • Silvicultural system: even-aged woods give clean boles; selection gives shorter, more tapering boles; coppice gives fuel and small timber.

5. Quick revision

  • Cell stages: division, enlargement and thickening, lignification. Pits link adjacent cells.
  • Heartwood is more durable because of toxic gums, resins and oils.
  • All-sapwood trees: fir, spruce, mango, haldu, semal, kadam.
  • Vessels are absent in conifers; tracheids do conduction and support.
  • Ring-porous: teak, toon (rare in India). Most Indian hardwoods are diffuse-porous.
  • Tyloses: improve durability, hinder impregnation.
  • Rays give silver grain; storeyed rays give ripple marks.

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